When Jak and Daxter receive a mysterious invite to Kras City for the reading of Krew's Last Will and Testament, they find they have been joined by familiar faces such as Ashelin, Keira, Samos and Torn. When Krew reveals in his recording will that everyone present, including his daughter, Rayn, has been poisoned, everyone is shocked. Krew gives everyone an ultimatum -- the only way to get the poison antidote in time is to race for him in the next Kras City Grand Championship, or they will all die.